*, *::before, *::after { box-sizing: border-box; }
html { scroll-behavior: smooth; }
body { margin: 0; color: var(--color-ink); background: var(--color-paper); font: 400 var(--step-0)/1.6 var(--font-body); text-rendering: optimizeLegibility; }
img { display: block; max-width: 100%; }
a { color: inherit; text-decoration-thickness: 1px; text-underline-offset: .22em; }
button, input { font: inherit; }
button { color: inherit; }
:focus-visible { outline: 3px solid var(--color-focus); outline-offset: 3px; }
.skip-link { position: fixed; z-index: 1000; top: .5rem; left: .5rem; transform: translateY(-160%); padding: .65rem .8rem; color: var(--color-ink); background: var(--color-surface); }
.skip-link:focus { transform: none; }

.site-shell { width: min(var(--page-max), 100%); margin-inline: auto; padding-inline: var(--page-gutter); }
.content-shell { width: min(var(--content-max), calc(100% - 2 * var(--page-gutter))); margin-inline: auto; }
.measure { max-width: var(--measure); }
.eyebrow { margin: 0 0 var(--space-4); font: 600 var(--step--1)/1.2 var(--font-body); letter-spacing: .12em; text-transform: uppercase; }
.display { margin: 0; font: 500 var(--step-5)/.87 var(--font-display); letter-spacing: -.055em; }
.section-title { margin: 0; font: 500 var(--step-4)/.98 var(--font-display); letter-spacing: -.04em; }
.subsection-title { margin: 0; font: 500 var(--step-2)/1.1 var(--font-display); }
.muted { color: var(--color-muted); }
.metadata { font: 500 var(--step--1)/1.45 var(--font-mono); }
.scientific-name { font-style: italic; }
.rule-top { border-top: var(--rule); }
.review-badge { display: inline-flex; gap: .4rem; align-items: center; padding: .28rem .48rem; border: 1px solid var(--color-review); border-radius: 99rem; color: #6f4610; background: var(--color-review-bg); font-size: var(--step--1); font-weight: 600; }

.button { display: inline-flex; min-height: 2.75rem; align-items: center; justify-content: center; padding: .65rem 1rem; border: 1px solid var(--color-ink); border-radius: var(--radius-small); color: var(--color-ink); background: transparent; font-weight: 600; text-decoration: none; cursor: pointer; transition: color var(--motion-fast), background var(--motion-fast), transform var(--motion-fast); }
.button:hover { color: var(--color-surface); background: var(--color-ink); }
.button:active { transform: translateY(1px); }
.button--forest { border-color: var(--color-forest); color: var(--color-surface); background: var(--color-forest); }
.button--forest:hover { background: var(--color-forest-dark); }
.button--quiet { border-color: var(--color-line); }

.site-header { position: sticky; z-index: 50; top: 0; display: grid; grid-template-columns: auto 1fr auto; align-items: center; min-height: 4.5rem; border-bottom: var(--rule); background: rgb(241 239 230 / .94); backdrop-filter: blur(12px); }
.brand { display: flex; gap: .65rem; align-items: baseline; font-weight: 700; text-decoration: none; }
.brand-mark { font-family: var(--font-display); font-size: 1.35rem; }
.primary-nav { display: flex; justify-content: center; gap: clamp(.8rem, 2.5vw, 2rem); }
.primary-nav a, .utility-nav a { font-size: .9rem; font-weight: 600; text-decoration: none; }
.utility-nav { display: flex; gap: 1rem; }
.menu-toggle { display: none; border: 1px solid var(--color-line); padding: .45rem .7rem; background: transparent; }

.photo-frame { margin: 0; }
.photo-frame img { width: 100%; object-fit: cover; }
.photo-frame figcaption { max-width: 70ch; padding-top: .7rem; color: var(--color-muted); font-size: var(--step--1); }
.photo-frame--wide img { aspect-ratio: 16 / 8; }
.photo-frame--documentary img { aspect-ratio: 4 / 3; }

.control-group { display: flex; flex-wrap: wrap; gap: .45rem; }
.control-group button { min-height: 2.6rem; padding: .5rem .7rem; border: 1px solid var(--color-line); border-radius: var(--radius-small); background: var(--color-surface); cursor: pointer; }
.control-group button[aria-pressed="true"] { border-color: var(--color-forest); color: var(--color-surface); background: var(--color-forest); box-shadow: inset 0 -3px 0 rgb(255 255 255 / .25); }
.control-group button:disabled { cursor: not-allowed; opacity: .5; }

.tooltip { position: absolute; z-index: 5; max-width: min(22rem, calc(100% - 2rem)); padding: .65rem .75rem; border: 1px solid rgb(255 255 255 / .5); border-radius: var(--radius-small); color: white; background: var(--color-overlay); box-shadow: var(--shadow-float); font-size: var(--step--1); pointer-events: none; }
.figure-note { margin: var(--space-4) 0 0; color: var(--color-muted); font-size: var(--step--1); line-height: 1.5; }
.source-links { display: flex; flex-wrap: wrap; gap: .8rem 1.2rem; margin-top: var(--space-5); font-size: var(--step--1); }

.site-footer { display: grid; grid-template-columns: 1fr auto; gap: var(--space-5); padding-block: var(--space-7); border-top: var(--rule); font-size: .88rem; }

@media (max-width: 58rem) {
  .site-header { grid-template-columns: 1fr auto; }
  .primary-nav, .utility-nav { display: none; }
  .menu-toggle { display: block; }
  .site-header.is-open .primary-nav { position: absolute; top: 100%; right: 0; left: 0; display: grid; gap: 0; padding: .7rem var(--page-gutter); border-bottom: var(--rule); background: var(--color-paper); }
  .site-header.is-open .primary-nav a { padding: .7rem 0; border-bottom: var(--rule); }
}

@media (max-width: 38rem) {
  .brand span:last-child { display: none; }
  .site-footer { grid-template-columns: 1fr; }
}

@media (prefers-reduced-motion: reduce) {
  html { scroll-behavior: auto; }
  *, *::before, *::after { scroll-behavior: auto !important; animation: none !important; transition-duration: 0ms !important; }
}